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To calculate the transferred score T i m from all taxonomic levels m to a protein pair from species i, we combine the scores S abk from orthologous groups consecutively from the lowest to the highest taxonomy level, subtracting the contributions from all lower taxonomic levels : where at each taxonomic level, we subtract the part of the score that originates from the species itself P ab i while additionally penalizing it for the number of paralogs in the respective orthologous groups f ab i and for the median self-normalized bit scores s a and s b of the proteins in the groups a and b.